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Bars indicate scaled ratios between the Doped and the Clean groups, with higher ratios of the Doped group to the left, and lower ratios to the right. The main findings of the study were that the doped athletes had higher lean mass, capillary density and myonuclei density, but lower maximal squat force relative to muscle mass and to fiber area, compared to the clean athletes. The Doped group also had a tendency towards larger fibers, although not significant, most likely due to large variations in fibre area. Low levels of LH and FSH, and high levels of prolactin in some individuals indicate a disturbed pituitary gland function with possible negative effects on reproductive function. High levels of ALAT, ASAT and CK in some individuals suggest that long term use of AAS could damage both liver and muscle tissue. However, no correlations between AAS intake and hormone levels was observed. Thus, testosterone levels at time of sampling cannot explain alternations in these variables, rather concentrations outside clinical limits must stem from long-term supplementation of AAS. Multivariate statistics showed that a combination of eight morphological parameters could clearly separate the doped from the clean athletes. Correlation analysis revealed significant positive correlations between AAS dosage and relative muscle force. The results support previous findings that AAS administration could induce enhancement in both muscle mass and muscle strength, and that the improvements are AAS dose-dependent [5], [8], [12], [25], [26]. This is the first study to confirm previous laboratory findings in active doping athletes. AAS and Muscle Mass, Muscle Morphology and Muscle Strength. Despite abundant studies on the effects of AAS on skeletal muscle, many results are contradictory [5]. Some studies have shown gains in body weight, girth, fat-free mass or lean body mass, but not in muscle strength [27]'[29], whereas others have shown gains in both muscle mass/girth and muscle strength [5], [8], [17], [30], [31], or in neither muscle mass/girth nor muscle strength after short term (from 17 days to 16 weeks) [32]'[35] or long term (2 years) [36] AAS administration. The present study showed that athletes with long term AAS supplementation had significantly higher leg lean mass compared to Clean athletes. The higher lean leg mass in the Doped subjects has been revealed to be mainly due to increased muscle mass, as shown in our previous study on the same group of doped athletes [37]. Increased muscle mass in subjects using AAS has been proposed to result from muscle hypertrophy alone [12] or from both muscle hypertrophy and hyperplasia [38]. Muscle hypertrophy is often evident by increased muscle fiber size and increased number of myonuclei. The latter is associated with satellite cell activation and myoblast infusion with the existing muscle fibers, leading to greater numbers of myonuclei in larger myofibers [39]. In previous studies on subjects with long term AAS supplementation (9’3. In the present study, long term AAS supplementation was only associated with higher lean leg mass, but not with larger fiber size, indicating that muscle fiber hyperplasia may play a role in the muscle mass enhancement, anabolic steroids buy aus. Coincidently, the number of myonuclei in type I fibers in the doped athletes was significantly higher than in the clean athletes, which may indicate satellite cell activation for muscle fiber hyperplasia. Not many studies have examined the effects of AAS on muscle capillaries. In a previous study of 20 weeks of graded testosterone enanthate injection (25, 50, 125, 300, or 600 mg), Sinha-Hikim et al. The authors suggested that it is not unlikely that a significant increase in capillaries takes longer than 20 weeks.
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